Description
Key Characters:
Growth Form:
Gynodioecious shrubs, 3–8 (–18) dm tall.
Stems:
Stems often swollen at the nodes, terete (cylindrical), 4–angled or –ridged, erect or strongly ascending, with few short lateral branches in the distal part of the stem, glabrous, internodes 0.5–5 cm long, green to greenish purple.
Roots:
Fibrous root system.
Leaves:
Leaves simple.
Opposite.
Blades narrowly elliptic to elliptic, sometimes the lower ones elliptic–oblanceolate or oblanceolate, 3–5 (–5.5) cm long, 0.6–1.5 cm wide.
Apex acute to acuminate.
Base leaf bases often connate around the stem, base attenuate to narrowly cuneate.
Blades thinly coriaceous, pale yellowish green, often slightly glaucous and reflective.
Margins entire, slightly thickened and weakly revolute.
1(–3) principal veins, the outer pair inconspicuous when present.
Petioles 0–0.3 cm long.
Stipules absent.
Flowers:
Flowers in inflorescence terminal, with 13–117 flowers, (4–) 8–23 cm long, usually laterally contracted, weakly congested, the hermaphroditic inflorescences slightly less congested and broader than the pistillate inflorescences; bracts of central axis 2–8 mm long, those of branches and flowers 1.5–3 mm long, green to purple–tinged, linear to subulate.
Flowers bisexual (perfect) or pistillate.
Calyx of hermaphroditic flowers with sepals 3.1–4.1 mm long, narrowly ovate, broadly and very shallowly navicular, oriented at 45° to 80° angle to the pedicel, green or tinged purple toward the apex, opaque, reflexed and convex at the base, producing a small transverse bulge, glabrous, margins scarious, sometimes somewhat irregularly undulate, entire, glabrous, apex sometimes inrolled, obtuse. Pistillate flowers: sepals 2.75–4.2 mm long, narrowly ovate, green or yellowish green, pale yellow at the base, opaque, reflexed and convex at the base, producing a small transverse bulge, broadly navicular to nearly flat, oriented at 60° to 90° angle to the pedicel, sometimes strongly upcurved near the middle, margins scarious and often irregularly undulate, glabrous, apex often inrolled, obtuse.
Corolla (petals) absent.
Stamens 10 (–12) in hermaphroditic flowers, sometimes a few (especially of the antisepalous whorl) non–functional and short, the filaments dimorphic, the antisepalous whorl 4–5.8 mm long, the alternate whorl 3.5–5 mm long; anthers 0.6–0.9 mm long, yellow. Pistillate flowers: Stamens 10 (–12), vestigial; filaments unequal, the antisepalous whorl 0.9–1.7 mm long, the alternate whorl (0.4–) 0.75–1 mm long; anthers 0.4–0.6 mm long, pale yellow, not producing pollen.
Ovary superior, 1-celled; ovule placentation free-central; styles (2–) 3 (–4), green (hermaphroditic flowers) or pale yellow (pistillate flowers).
Fruit:
Capsules 4.5–5 mm long; ovoid.
Seeds 0.7–0.9 mm long; orbicular–reniform; the surface papillose; papillae of the margins larger than those of the faces; perisperm starchy; hard or rarely soft; true endosperm absent.
Ploidy:
2n = 55*–60*; probably 60*
Habitat:
Scattered on ridges and steep slopes usually beyond the reach of grazing animals in remnant dry shrubland.
Elevation Range:
180–670 m.